Stefan Lohmer, Massimo Maddaloni, Mario Motto, Francesco Salamini, Richard D. Thompson, Translation of the mRNA of the Maize Transcriptional Activator Opaque-2 Is Inhibited by Upstream Open Reading Frames Present in the Leader Sequence, The Plant Cell, Vol. 5, No. 1 (Jan., 1993), pp. 65-73
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Lohmer et al. (1993) studied this question.