Richard Etheridge, A Review of Lizards of the Liolaemus wiegmannii Group (Squamata, Iguania, Tropiduridae), and a History of Morphological Change in the Sand-Dwelling Species, Herpetological Monographs, Vol. 14 (2000), pp. 293-352
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Richard Etheridge (2000) studied this question.
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