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Frank Seebacher, Ross A. Alford, Movement and Microhabitat Use of a Terrestrial Amphibian (Bufo marinus) on a Tropical Island: Seasonal Variation and Environmental Correlates, Journal of Herpetology, Vol. 33, No. 2 (Jun., 1999), pp. 208-214
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Seebacher et al. (1999) studied this question.
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