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The hypothesis of a locomotor role for the free rays of Trigloporus lastoviza is supported by: (1) their periodic retraction and protraction during slow displacement over the substratum whenever the propulsive body wave responsible for swimming is not visible; (2) the integration of the rays' individual cycles into a hexapod gait apparently generated by a metachronal impulse; and (3) the kinematic features of the longest free ray in relation to the bottom-walking velocity.
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Renous et al. (2000) studied this question.
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