Competition studies in continuous culture between a Hup− mutant of Azotobacter chroococcum and its presumed isogenic Hup+ recombinant showed that Hup activity benefited the organism under N2-fixing, sucrose- or phosphate-limiting conditions but was ineffective or disadvantageous under O2-, sulphate- or iron-limitation.
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Yates et al. (1989) studied this question.
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