Kuniaki Watanabe, Robert M. King, Tetsukazu Yahara, Motomi Ito, Jun Yokoyama, Takeshi Suzuki, Daniel J. Crawford, Chromosomal Cytology and Evolution in Eupatorieae (Asteraceae), Annals of the Missouri Botanical Garden, Vol. 82, No. 4 (1995), pp. 581-592
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Watanabe et al. (1995) studied this question.