Geffery R. Luttrell, Anthony A. Echelle, William L. Fisher, David J. Eisenhour, Declining Status of Two Species of the Macrhybopsis aestivalis Complex (Teleostei: Cyprinidae) in the Arkansas River Basin and Related Effects of Reservoirs as Barriers to Dispersal, Copeia, Vol. 1999, No. 4 (Dec. 17, 1999), pp. 981-989
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Luttrell et al. (1999) studied this question.