Wood Anatomy of Onagraceae: Additional Species and Concepts

vod anatomy of Epilobium colchicum subsp.ag je eeu excorticata, and Hauya heydean is described qualitatively and quantitatively.For the latter two species, large logs vere available and wood portions from both inside and 1 were analyzed.Although these | e species offer no features new for Onagraceae, each adds features new for its respective genus.By means of numerical indices which are termed vulnerability and ABER respectively, values are presented to show the range in ecological d of woods of the three species, as well as of all Onagraceae studied earlier.Onagraceae d. a wide range in these indices and probably form a good model of what use indie in families with a broad ecological range will demonstrate.Wood from inside of logs of Fuchsia excorticata and Hauya heydeana is more xeromorphic than wood from the periphery.In my monograph of wood anatomy of Onagraceae (Carlquist, 1975a), I attempted a sampling of woods based largely on availability of portions of suitable size.In any family in which herbs predominate, one is faced with arbitrary decisions as to whether some species which form little secondary xylem should be included.However, after the appearance of the monograph, Dr. Peter H Raven placed at my disposal three wood samples with abundant secondary xylem for their respective genera, and otherwise cf more than passing interest. MATERIALS AND METHODSEpilobium colchicum Alboff subsp.colchicum (sect.Chamaenerion) was collected by Dr. Peter H. Raven from a streambed near the Lagodekhi Hotel in eastern Georgia, U.S.S.R., and is unusually woody for an Epilobium.The species of Fuchsia that forms perhaps the largest trees in that genus (and perhaps also in the family) is F. excorticata (J.R. & G. Forst.)L. f.A section from the base of a tree approximately 0.9 m in diameter (outline of section irregular) was supplied by the New Zealand Institute of Forestry.Dr. Dennis E. Breedlove's mesoamerican field work yielded a log, approximately 25 cm in diameter, of Hauya heydeana Donnell Smith.Because all the Hauya wood samples utilized in my earlier study were of H. elegans subsp.cornuta, material of the second species of this interesting genus was especially welcome.All three wood samples were dried.Methods of study were the same as those employed for dried samples in the earlier paper.Dr. Larry DeBuhr's work in preparing sections and macerations and in obtaining data is gratefully acknowledged.Because differences in wood anatomy were observed (Carlquist, 1975a) in samples of different diameter in Hauya elegans subsp.cornuta, both inner and outer portions of 1 This study has been aided by a grant from the National Science Foundation, BMS 73-07055-Al.The wood of Fuchsia excorticata was provided by Dr. E.