Didier Cabrillac, , Jerome Garin, , , Christian Dumas, Thierry Gaude, J. Mark Cock, The S₁₅ Self-Incompatibility Haplotype in Brassica Oleracea Includes Three S Gene Family Members Expressed in Stigmas, The Plant Cell, Vol. 11, No. 5 (May, 1999), pp. 971-986
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Cabrillac et al. (1999) studied this question.