Peter H. Thrall, Andrew M. Jarosz, Host-Pathogen Dynamics in Experimental Populations of Silene Alba and Ustilago Violacea. I. Ecological and Genetic Determinants of Disease Spread, Journal of Ecology, Vol. 82, No. 3 (Sep., 1994), pp. 549-559
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Thrall et al. (1994) studied this question.