Hongyong Fu, Soo Young Kim, William D. Park, High-Level Tuber Expression and Sucrose Inducibility of a Potato Sus4 Sucrose Synthase Gene Require 5′ and 3′ Flanking Sequences and the Leader Intron, The Plant Cell, Vol. 7, No. 9 (Sep., 1995), pp. 1387-1394
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Fu et al. (1995) studied this question.