M. Carlsson, L. M. Nilsson, Bj. Svensson, S. Ulfstrand, R. S. Wotton, Lacustrine Seston and Other Factors Influencing the Blackflies (Diptera: Simuliidae) Inhabiting Lake Outlets in Swedish Lapland, Oikos, Vol. 29, No. 2 (1977), pp. 229-238
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Carlsson et al. (1977) studied this question.
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