James E. Cresswell, Spatial Heterogeneity, Pollinator Behaviour and Pollinator-Mediated Gene Flow: Bumblebee Movements in Variously Aggregated Rows of Oil-Seed Rape, Oikos, Vol. 78, No. 3 (Apr., 1997), pp. 546-556
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James Cresswell (1997) studied this question.
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