R. K. Misra, Vectorial Analysis for Genetic Clines in Body Dimensions in Populations of Drosophila subobscura Coll. and a Comparison with Those of D. robusta Sturt., Biometrics, Vol. 22, No. 3 (Sep., 1966), pp. 469-487
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R. K. Misra (1966) studied this question.
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