Lyle P. Brinton, James H. Oliver, Jr., Gross Anatomical, Histological, and Cytological Aspects of Ovarian Development in Dermacentor andersoni Stiles (Acari: Ixodidae), The Journal of Parasitology, Vol. 57, No. 4 (Aug., 1971), pp. 708-719
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Brinton et al. (1971) studied this question.
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