Using a collection of liquid-preserved vegetative shoots aa all 17 genera of а I investigated the cross-sectional histology of mature leaves.The study focussed оп a search systematically useful and diagnostic features as well as on the determination of Seele trends of specialization of leaf features.Exclusively dorsiventral-leaved rad are found in Ludwi igia, Hauya, Fuchsia, Circaea, and Lopezia.Leaves of these genera also have the most prominent structure.The most generalized anatomy including the presence of exraylary aes and diverse calcium oxalate crystal structure i is found i in Fuchsia, Ludwi igia, and Hauya.The other twelve genera, which belong with data from wood anatomy, pollen morphology, reproductive biology, and biogeography, I hyмлан that the Onagraceae leaves show a general reduction in leaf complexity.Leaf anatomy by itself is not diagnostic for those genera with reduced structure but it may be distinctive in the genera with more complex structure such as Fuchsia, Ludwigia, and Hauya.
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Richard C. Keating (1982) studied this question.