In 1966, Ueno described and figured a nymph as Ephemerella sp. from a series of 12 specimens collected by the Kyoto University Pamir–Hindukush Expedition in 1955. Allen (1971) named this mayfly E. borakensis from Ueno's description and figure, and placed the species in the subgenus Drunella Needham.
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Richard K. Allen (1974) studied this question.
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