Michael N. Manlove, Ramone Baccus, Michael R. Pelton, Michael H. Smith, David Graber, Biochemical Variation in the Black Bear, Bears: Their Biology and Management, Vol. 4, A Selection of Papers from the Fourth International Conference on Bear Research and Management, Kalispell, Montana, USA, February 1977 (1980), pp. 37-41
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Manlove et al. (1980) studied this question.
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