J. Mallet, J. T. Longino, D. Murawski, A. Murawski, A. Simpson De Gamboa, Handling Effects in Heliconius: Where Do All the Butterflies Go?, Journal of Animal Ecology, Vol. 56, No. 2 (Jun., 1987), pp. 377-386
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Mallet et al. (1987) studied this question.
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