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Charles C. Rhoades, Robert L. Sanford, Jr., David B. Clark, Gender Dependent Influences on Soil Phosphorus by the Dioecious Lowland Tropical Tree Simarouba amara, Biotropica, Vol. 26, No. 4 (Dec., 1994), pp. 362-368
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Rhoades et al. (1994) studied this question.
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