The Production of Dry Matter by Swards of Perennial Ryegrass, Differing in Average Age of Foliage

Leaves of different ages were removed 7 times between June and October. Each defoliation was timed to match the appearance of successive leaves on each tiller. The removal of leaves at progressively younger stages reduced total production above ground, but the most severe treatment, the removal of all expanded leaves repeatedly, still produced about 75% as much as the least severely defoliated sward. Clipping with shears 3 times reduced yield as much as removing all expanded leaves 7 times. A crop allowed to grow untouched apparently produced more than any defoliated treatment, but this may have been an artefact. The lower production following increasingly drastic defoliation was not associated with reduced concentrations of nitrogen or potassium in the foliage removed, or in the crop left behind. The total quantity of these elements recovered in the crop was greatest in those treatments where least dry matter was produced. Explanations for the results depend on the capacity of swards of predominantly young leaves to assimilate carbon more rapidly than those containing old and senescent tissue.