Brent D. Mishler, Peter H. Thrall, John S. Hopple, Jr., Efrain De Luna, Rytas Vilgalys, A Molecular Approach to the Phylogeny of Bryophytes: Cladistic Analysis of Chloroplast-Encoded 16S and 23S Ribosomal RNA Genes, The Bryologist, Vol. 95, No. 2 (Summer, 1992), pp. 172-180
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Mishler et al. (1992) studied this question.