Anaspidacea, Bathynellacea (Crustacea, Syncarida), generalised tracks, and the biogeographical relationships of South America

The spatial evolution of South American Syncarida (Anaspidacea and Bathynellacea) and related taxa is evaluated applying a panbiogeographic approach, where Ocean basins are used to identify major patterns of intercontinental distribution. The Pacific basin, corresponding to a southern temperate track, is identified as the major evolutionary centre for Stygocaris(Stygocarididae), Bathynella (Bathynellidae), Atopobathynellu, and Chilibathynella (Parabathynellidae), whereas distribution of Nannobathynella (Bathynellidae) and Cteniobathynella (Parabathynellidae) is centred on the Atlantic Ocean, belonging to a northern tropical track. It is concluded that the biotic origin of the South American Syncarida is complex, implying the existence of at least two ancestral biotas.