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Randall J. Bayer, Patterns of Isozyme Variation in Western North American Antennaria (Asteraceae: Inuleae). I. Sexual Species of sect. Dioicae, Systematic Botany, Vol. 13, No. 4 (Oct. - Dec., 1988), pp. 525-537
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Randall J. Bayer (1988) studied this question.
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