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A new genusPusulissusgen. nov.with two new speciesP. phiaoacensissp. nov.andP. coronomensissp. nov.are described respectively from Vietnam and China in the tribe Parahiraciini of the family Issidae. A molecular phylogeny using combined genes (18S, 28S, COX1 and Cytb) confirms its placement in the tribe Parahiraciini. The tribe is divided into three subtribes: Scantiniinasubtribe nov.with the type genusScantiniusStål, 1866, and Vindilisinasubtribe nov.with type genusVindilisStål, 1870 plusNisoprincessaGnezdilov, 2017, and the nominal subtribe Parahiraciina Cheng & Yang, 1991subtribe nov.The characteristics of these subtribes are provided, with a key to identification. GenusFolifemurumChe, Zhang & Wang, 2013 is transferred to HemisphaeriiniMongolianina, and genusGelastyrellaYang, 1994 is maintained as a valid name.
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Bourgoin et al. (2020) studied this question.
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