Euoniticellus intermedius (Coleoptera: Scarabaeidae): Description of Adults and Immatures and Biology of Adults

The larva and pupa of Euoniticellus intermedius (Reiche) are described and illustrated. Nesting behavior conforms to Halffter and Edmonds (1982, Publ. 10, Instituto de Ecologia, Mexico, D.F.) Pattern I; an egg is laid in a brood mass constructed at the end of a burrow. The egg chamber is much larger than that described for other species and is located centrally in the mass. Temperature is demonstrated to have a direct impact on longevity and fecundity of adults and upon developmental time of immatures.