In this study, we demonstrated that the putative Vibrio fischeri rpoN gene, which encodes sigma(54), controls flagellar biogenesis, biofilm development, and bioluminescence. We also show that rpoN plays a requisite role initiating the symbiotic association of V. fischeri with juveniles of the squid Euprymna scolopes.
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Wolfe et al. (2004) studied this question.