Arkady B. Rovinsky, Hendra Adiwidjaja, Vladimir Z. Yakhnin, Michael Menzinger, Patchiness and Enhancement of Productivity in Plankton Ecosystems due to the Differential Advection of Predator and Prey, Oikos, Vol. 78, No. 1 (Feb., 1997), pp. 101-106
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Rovinsky et al. (1997) studied this question.
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