T. F. Dougherty, B. J. Stover, J. H. Dougherty, W. S. S. Jee, C. W. Mays, C. E. Rehfeld, W. R. Christensen, H. C. Goldthorpe, Studies of the Biological Effects of Ra226, Pu²³⁹, Ra²²⁸( MsTh₁), Th²²⁸( RdTh), and Sr⁹⁰ in Adult Beagles, Radiation Research, Vol. 17, No. 4 (Oct., 1962), pp. 625-681
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Dougherty et al. (1962) studied this question.