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Changes in population density were evaluated over an outbreak cycle of the Douglas-fir tussock moth. Growth rate of populations was independent of larval density in the first year, but thereafter became a reciprocal function of density, terminated by complete population collapse at the end of the third year. Through correlation analyses, it was found that density of small larvae accounted for a large proportion of the change in population density between years. This suggests that the local infestations developed largely from resident populations of at least 2 years and not from spread of early instar larvae during the outbreak.
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Richard R. Mason (1974) studied this question.
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