High-speed films of sound-producing male Lycosa and Schizocosa spp. show that their palps are not "drummed." as previously assumed. but rapidly oscillate at the tibio-tarsal joint. A new type of spider stridulatory organ is located on hidden surfaces of this joint. Coupling to the substratum via tarsal spines increases the communicatory effectiveness of this apparatus.
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Jerome S. Rovner (1975) studied this question.