The dynamic effects of spatial heterogeneity are discussed in relation to three systems: (1) a single-species system, (2) a parasitoid-host system, and (3) a disease-host system. In each case, a heterogeneous environment of discrete patches is specified, between which the animals are distributed. The central conclusion is the same irrespective of the system: the more contagious the distribution of the animals per patch, the greater is the population stability. In the single-species and disease-host systems, this arises because increased clumping accentuates the density dependent mortality within a patch, while in the parasitoid-host system it results from differential susceptibility of host individuals to parasitism.
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M. P. Hassell (1980) studied this question.
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