Timothy W. McNellis, Albrecht G. von Arnim, Takashi Araki, Yoshibumi Komeda, , Xing-Wang Deng, Genetic and Molecular Analysis of an Allelic Series of cop1 Mutants Suggests Functional Roles for the Multiple Protein Domains, The Plant Cell, Vol. 6, No. 4 (Apr., 1994), pp. 487-500
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McNellis et al. (1994) studied this question.