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Mitsuyasu Hasebe, Paul G. Wolf, Kathleen M. Pryer, Kunihiko Ueda, Motomi Ito, Ryosuke Sano, Gerald J. Gastony, Jun Yokoyama, James R. Manhart, Noriaki Murakami, Edmund H. Crane, Christopher H. Haufler, Warren D. Hauk, Fern Phylogeny Based on rbcL Nucleotide Sequences, American Fern Journal, Vol. 85, No. 4, Use of Molecular Data in Evolutionary Studies of Pteridophytes (Oct. - Dec., 1995), pp. 134-181
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Hasebe et al. (1995) studied this question.
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