Olusegun O. Osunkoya, Julian E. Ash, Mike S. Hopkins, Andrew W. Graham, Influence of Seed Size and Seedling Ecological Attributes on Shade-Tolerance of Rain-Forest Tree Species in Northern Queensland, Journal of Ecology, Vol. 82, No. 1 (Mar., 1994), pp. 149-163
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Osunkoya et al. (1994) studied this question.
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