Marilyn A. Anderson, Geoffrey I. McFadden, Robert Bernatzky, Angela Atkinson, Timothy Orpin, Helen Dedman, Geoffrey Tregear, Ross Fernley, Adrienne E. Clarke, Sequence Variability of Three Alleles of the Self-Incompatibility Gene of Nicotiana alata, The Plant Cell, Vol. 1, No. 5 (May, 1989), pp. 483-491
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Anderson et al. (1989) studied this question.