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G. R. Miller, Adam Watson, Territories and the Food Plant of Individual Red Grouse: I. Territory Size, Number of Mates and Brood Size Compared with the Abundance, Production and Diversity of Heather, Journal of Animal Ecology, Vol. 47, No. 1 (Feb., 1978), pp. 293-305
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Miller et al. (1978) studied this question.
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