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Christer Wiklund, Johan Forsberg, Sexual Size Dimorphism in Relation to Female Polygamy and Protandry in Butterflies: A Comparative Study of Swedish Pieridae and Satyridae, Oikos, Vol. 60, No. 3 (Apr., 1991), pp. 373-381
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Wiklund et al. (1991) studied this question.
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