James M. Cooper, Jerry Wilhm, Spatial and Temporal Variation in Productivity, Species Diversity, and Pigment Diversity of Periphyton in a Stream Receiving Domestic and Oil Refinery Effluents, The Southwestern Naturalist, Vol. 19, No. 4 (Jan. 20, 1975), pp. 413-427
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Cooper et al. (1975) studied this question.