Timothy R. Schmidt, Joseph P. Bielawski, John R. Gold, Molecular Phylogenetics and Evolution of the Cytochrome b Gene in the Cyprinid Genus Lythrurus (Actinopterygii: Cypriniformes), Copeia, Vol. 1998, No. 1 (Feb. 3, 1998), pp. 14-22
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Schmidt et al. (1998) studied this question.
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