Stefan Eberhard, Nancy Doubrava, , Debra Mohnen, Audrey Southwick, Alan Darvill, Peter Albersheim, Pectic Cell Wall Fragments Regulate Tobacco Thin-Cell-Layer Explant Morphogenesis, The Plant Cell, Vol. 1, No. 8 (Aug., 1989), pp. 747-755
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Eberhard et al. (1989) studied this question.