T. F. Mace, C. C. Davis, Energetics of a Host-Parasite Relationship as Illustrated by the Leech Malmiana Nuda, and the Shorthorn Sculpin Myoxocephalus Scorpius, Oikos, Vol. 23, No. 3 (1972), pp. 336-343
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Mace et al. (1972) studied this question.
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