T. T. Veblen, D. H. Ashton, F. M. Schlegel, A. T. Veblen, Distribution and Dominance of Species in the Understorey of a Mixed Evergreen-Deciduous Nothofagus Forest in South-Central Chile, Journal of Ecology, Vol. 65, No. 3 (Nov., 1977), pp. 815-830
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Veblen et al. (1977) studied this question.
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