Past studies on titmice have found song to serve exclusively in defense of territory (e.g., Dixon 1949, Lemon 1968.Krebs et al. 1978).vet Krebs et al. (1981) showed I_ .that in the Great Tit (Purus major) song ma; also' function to attract females.They noted that after a female was removed from a mated pair, the male increased song output almost six-fold and, when reunited with the female, subsequently decreased his singing.The study population was located 22 km N of Flagstaff, Arizona in a pinyon-juniper-ponderosa pine ecotone.In 1982.five males and some females were marked with U.S. Fish ind Wildlife Service aluminum bands and unique combinations of colored plastic leg-bands.Birds were observed between 06:OO and 12:OO usuallv for a ueriod of one hour per day several times a week.Observations were made from early March through late May.Songs, behavioral observations, and timing (from running stopwatches) were recorded simultaneously on a Uher 4000 Report IC tape recorder with a Dan Gibson parabolic microphone.While studying the singing behavior of the Plain Tit-This report deals with the behavior of four territorial males, all of whom were mated at the beginning of the JEHL, J. R., JR. 1974.The near-shore avifauna of the Middle American west coast.Auk 9 1:68 l-699.
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L. Scott Johnson (1983) studied this question.