Brita M. Svensson, Bengt A. Carlsson, P. Staffan Karlsson, K. Olle Nordell, Comparative Long-Term Demography of Three Species of Pinguicula, Journal of Ecology, Vol. 81, No. 4 (Dec., 1993), pp. 635-645
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Svensson et al. (1993) studied this question.
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