Bruce E. Mahall, Roderic B. Park, The Ecotone Between Spartina Foliosa Trin. and Salicornia Virginica L. in Salt Marshes of Northern San Francisco Bay: III. Soil Aeration and Tidal Immersion, Journal of Ecology, Vol. 64, No. 3 (Nov., 1976), pp. 811-819
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Mahall et al. (1976) studied this question.
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