Sylvia J. Lilly, J. F. Sloane, R. Bassindale, F. J. Ebling, J. A. Kitching, The Ecology of the Lough Ine Rapids with Special Reference to Water Currents, Journal of Animal Ecology, Vol. 22, No. 1 (May, 1953), pp. 87-122
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Lilly et al. (1953) studied this question.