Samuel C. Zeeman, Takayuki Umemoto, Wei-Ling Lue, Pui Au-Yeung, Cathie Martin, Alison M. Smith, Jychian Chen, A Mutant of Arabidopsis Lacking a Chloroplastic Isoamylase Accumulates Both Starch and Phytoglycogen, The Plant Cell, Vol. 10, No. 10 (Oct., 1998), pp. 1699-1711
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Zeeman et al. (1998) studied this question.