Canonical correspondence analysis (CCA) was used to examine the distribution of leeches (Annelida: Hirudinea) in 18 lakes at the Experimental Lakes Area (ELA). Leech community composition was best described by an ordination incorporating alkalinity, primary productivity, and lake area. In general, highest species richness occurred in small, eutrophic lakes whereas lowest richness was recorded in medium to large lakes with low productivity. Contrary to results for some other taxa, lake pH was not a dominant variable, describing only a small amount of variance in the species–environment relationship.
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Grantham et al. (1994) studied this question.
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