M. A. Baars, Th. S. Van Dijk, Population Dynamics of Two Carabid Beetles at a Dutch Heathland: I. Subpopulation Fluctuations in Relation to Weather and Dispersal, Journal of Animal Ecology, Vol. 53, No. 2 (Jun., 1984), pp. 375-388
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Baars et al. (1984) studied this question.
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